Nikon's lightweight S-line 400mm prime for Z-mount — VR stabilised, compact for its class, and optically superb.
The Nikon Nikkor Z 400mm f/4.5 VR S was released in 2022 as a compact super-telephoto prime for Nikon Z-mount, providing 400mm at f/4.5 in a significantly lighter package than traditional 400mm f/4 and f/2.8 super-telephotos. S-line designation. 5.5-stop VR. At 1,245g with tripod collar and 95mm filter thread. Minimum focus distance of 2.5m. Compatible with Z 1.4x and 2x teleconverters.
The optical design uses 19 elements in 13 groups including 1 ED, 2 Super ED, and 1 SR (short wavelength refractive) element. Nine aperture blades. The 95mm filter thread. At 1,245g with tripod collar the lens is moderate for a 400mm prime. Minimum focus distance of 2.5m. 5.5-stop VR. Stepping motor AF. S-line. Compatible with Z 1.4x and 2x teleconverters.
The 1,245g weight makes the Z 400mm f/4.5 VR S substantially lighter than 400mm f/4 and f/2.8 alternatives — lighter than the Nikon AF-S 500mm f/4E FL ED VR (3090g) and competitive with 500mm f/5.6 PF designs. The SR element enables the weight reduction by providing the short wavelength refractive power that traditionally requires large ED element groups. The 5.5-stop VR extends handheld wildlife telephoto capability.
On the used market the Nikon Z 400mm f/4.5 VR S is available at professional telephoto prime pricing. Condition checks: VR engagement, stepping motor AF response, 95mm front element for marks, and tripod collar rotation. The 2.5m MFD limits very close wildlife approach. Compatible with all Nikon Z-mount FX bodies; Z 1.4x TC extends to 560mm f/6.3.